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holds grant NIH/NIAAA (5R24AA022057)

3.3 Melanin-synthesis-inhibiting peptides of mushrooms and bacteria In a 1974 study, a peptide (Madhosingh and Sundberg, 1974) with tyrosinase inhibitory activity was first reported

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Metformin enhances mitochondrial biogenesis and thermogenesis in brown adipocytes of mice

Additionally, another transcription factor, AP-1 (activating protein-1) is significantly involved in the cellular stress response of melanocytes

Materials and methods Experimental animals The animal experiments were performed on the basis of ethical regulations for animal testing, and the animal protocol employed in this study was approved by the Animal Experimentation Committee of Jinan University (IACUC-20191212-02)