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In contrast, an increased spermidine/spermine ratio (resulting from a spermine synthase gene mutation in mice) was linked to increased anxiety, downregulated mitochondrial oxidative phosphorylation and ribosomal protein synthesis in the cerebral cortex, as well as impaired mitochondrial bioenergetics in isolated primary fibroblasts (Akinyele et al

It features an Arg3 substitution and an N-terminal extension that researchers study for altered receptor binding and reduced interaction with IGF binding proteins

Complex I (NADH-ubiquinone oxidoreductase) activity was assayed by measuring decreased absorbance at 340 nm, indicating reduction of NADH oxidation