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Interestingly, some PAHs, such as benzo(a)pyrene, have been reported to induce necrosis through activation of PARP-1 (Lin et al., 2008), whereas nicotine has been reported to down-regulate the expression of PARP-1 (Lu et al., 2017), reinforcing the hypothesis that, even when present in lower doses compared to tobacco smoke, PAHs in HnB mainstream smoke can contribute to testicular toxicity

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doi: 10.1089/ars.2011.4475 20 CaoSSKaufmanRJ

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